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It depends on where it's at. Most common use is in a LIKE statement, it is a single character wild card.
ex; LIKE = ???toon will give you all things that has toon with three letters before it such as cartoon or flytoon or bumtoon. See also the * wild card and the %.
I know it appears in the WHERE statement as a variable placeholder that will be filled in, but don't know how that works.
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